(Carcharhinidae)

Requiem Sharks

Сірі акули

The requiem shark family (Carcharhinidae) is comprised of 11 genera and at least 56 described species worldwide. This is one of the largest, most important shark families, and includes many common and wide-ranging species. These sharks are dominant (in terms of their biodiversity, abundance and biomass) in tropical continental shelf and offshore habitats, but some species are also found in subtropical and warm-temperate seas. Several members of this family are closely associated with coral reefs and oceanic islands, while other species range far into ocean basins. One pelagic requiem shark, the Blue Shark, has one of the greatest geographic ranges of any shark or marine vertebrate, from high latitude, cool-temperate waters into the tropics. A few other species are found in temperate waters and some even at great depths, but none are truly specialised deepsea sharks (compared with many species of the families Squalidae and Pentanchidae). Some requiem shark species occur in freshwater rivers and lakes. Although members of other families may enter river-mouths and ascend rivers for a short distance, the little-known Central Indo-Pacific river sharks, Glyphis species, and the broadly distributed Bull Shark appear to be the only living sharks that can live in fresh water for extended periods. The Bull Shark has a wide range in tropical and warm-temperate rivers and lakes around the world and is remarkable for the apparent ease with which it can move from saline to freshwater conditions and back again.

Although some species are relatively small (65-100cm), most are medium to large in size, with maximum lengths ranging to nearly 400cm. They have a long, arched mouth with blade-like teeth (often broader in the upper jaw), most with short labial furrows (except Rhizoprionodon), no nasoral grooves or barbels. Usually round (to horizontal) eyes with internal nictitating eyelids, usually no spiracles. Two dorsal fins, one anal fin, first dorsal fin medium to large with base well ahead of pelvic bases, second usually much smaller. Precaudal pits, caudal fin with a strong lower lobe and lateral undulations on upper margin. Mostly unpatterned (particularly in Carcharhinus). The extremely rare river sharks (Glyphis species), the only truly freshwater shark species, are very difficult to distinguish without tooth and vertebral counts.

All requiem sharks are viviparous with a yolk-sac placenta and have litters ranging in size from just one or two pups to 135 in the Blue Shark. They are strong, active swimmers, occurring singly or in small to large schools. Some species are ‘ram-ventilators’ needing to swim continually to oxygenate their gills, while others are capable of resting motionless for extended periods on the bottom. Some are solitary or socialise in small groups, some are social schooling species. There is a clear hierarchical dominance between certain species: Oceanic Whitetip Sharks are dominant over Silky Sharks of the same size, which in turn can dominate Grey Reef Sharks; Galapagos Sharks are dominant over Blacktip Sharks but subordinate to the Silvertip.

Requiem sharks are extraordinarily fast and effective hunters. Their elongated, torpedo-shaped bodies make them quick and agile swimmers, so they can easily attack any prey. Some species are continually active, while others are capable of resting motionless for extended periods on the bottom. They have a range of food sources depending on location and species, including bony fish, squid, octopus, lobster, turtles, marine mammals, seabirds, other sharks and rays; smaller species tend to select a narrow range of prey, but some very large species are virtually omnivorous. They are often considered the “garbage cans” of the seas because they will eat almost anything, even non-food items like trash. They are migratory hunters that follow their food source across entire oceans. They tend to be most active more active at night, or dawn and dusk, where their impressive eyesight can help them sneak up on unsuspecting prey. Most requiem sharks hunt alone, however some species like the whitetip reef sharks and lemon sharks are cooperative feeders and will hunt in packs through coordinated, timed attacks against their prey. Some of the species have been shown to give specialized displays when confronted by divers or other sharks, which may be indicative of aggressive or defensive threat.

Bronze Whaler, Spinner Shark, Silky Shark, Bull Shark, Dusky Shark, Galápagos Shark, Common Blacktip Shark, Oceanic Whitetip Shark, Blacktip Reef Shark, Caribbean Reef Shark, Sandbar Shark, Blue Shark, Lemon Shark, Sicklefin Lemon Shark, Whitetip Reef Shark, Grey Reef Shark, Graceful Shark are considered dangerous or potentially dangerous to humans.

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carcharhinus acronotus

(Carcharhinus acronotus)

Blacknose Shark

Акула чорноноса

Total length: 1.3–2 m.
Weight: 10–18.9 kg.

It inhabits the continental and insular shelves off the eastern coast of the Americas, as far north as North Carolina and as far south as southern Uruguay, including the Bahamas, the Gulf of Mexico, and the Caribbean Sea. It occurs at depths of 9–64 m.

carcharhinus albimarginatus

(Carcharhinus albimarginatus)

Silvertip Shark

Акула срібнопера

Total length: 2–3 m.
Weight: <136 kg.

It has a fragmented distribution throughout the tropical Indian and Pacific Oceans. It occurs over continental and insular shelves at depths of 30–800 m and is most common around isolated islands, coral banks, and reef drop-offs.

carcharhinus altimus

(Carcharhinus altimus)

Bignose Shark

Акула великоноса

Total length: 2.7–2.8 m.

It is distributed worldwide in tropical and subtropical waters, this migratory shark frequents deep waters around the edges of the continental shelf. It is typically found at depths of 90–430 m.

carcharhinus amblyrhynchoides

(Carcharhinus amblyrhynchoides)

Graceful Shark

Сіра акула витончена

Total length: <1.7 m.

It is widely distributed in the tropical Indo-Pacific, with records from the Gulf of Aden, southwestern India and Sri Lanka, the Gulf of Thailand, Vietnam, the Philippines, Borneo and Java, and from Papua New Guinea to northern Australia. It is an open-water inhabitant that can be found from close to shore to the outer continental and insular shelves, diving at least 50 m.

carcharhinus amblyrhynchos

(Carcharhinus amblyrhynchos)

Grey Reef Shark

Сіра рифова акула

Total length: 1.8–2.6 m.
Weight: <33.7 kg.

It is found in the Indo-Pacific, ranging from South Africa in the west to Easter Island in the east. It is most often seen in shallow waters near coral reef drop-offs at depths of less than 60 m, but it may occasionally descend to depths of up to 1,000 m.

carcharhinus amboinensis

(Carcharhinus amboinensis)

Pigeye Shark

Свиноока акула

Total length: 1.9–2.8 m.

It is found in warm coastal waters of the eastern Atlantic and western Indo-Pacific, at depths from the surface to 150 m.  It is distributed from Nigeria in the Gulf of Guinea to Eritrea in the Red Sea, and it occasionally reaches the Mediterranean Sea. It also occurs from the southern Arabian Peninsula eastward to Indonesia, Indochina, the Philippines, southern China, New Guinea, and northern and eastern Australia.

carcharhinus borneensis

(Carcharhinus borneensis)

Borneo Shark

Сіра акула борнеоська

Total length: <70 cm.

It is known only from inshore waters around Mukah in north-western Borneo and Zhoushan Island in China, though it may once have been more widely distributed.

carcharhinus brachyurus

(Carcharhinus brachyurus)

Bronze Whaler

Акула бронзова

Total length: <3.2 m.
Weight: <305 kg.

It is distributed in a number of separate populations in the northeastern and southwestern Atlantic, off southern Africa, in the northwestern and eastern Pacific, and around Australia and New Zealand, with scattered reports from equatorial regions. The species can be found from brackish rivers and estuaries to shallow bays and harbors, to offshore waters 100 m deep or more.

carcharhinus brevipinna

(Carcharhinus brevipinna)

Spinner Shark

Сіра акула короткоплавцева

Total length: 2–3 m.
Weight: 50–90 kg.

It occurs in tropical and warm temperate waters worldwide, except for in the eastern Pacific Ocean. It has been reported from the ocean surface to a depth of 100 m, though it prefers water less than 30 m deep. It may be found from coastal waters to well offshore, over continental and insular shelves.

carcharhinus cautus

(Carcharhinus cautus)

Nervous Shark

Сіра акула нервова

Total length: 1–1.5 m.

It is found over continental and insular shelves off northern Australia from Shark Bay in the west to Moreton Bay in the east, as well as off Papua New Guinea and around the Solomon Islands. It generally inhabits shallow inshore waters, at depths from the surface to 20 m.

carcharhinus cerdale

(Carcharhinus cerdale)

Pacific Smalltail Shark

Тихоокеанська дрібнохвоста акула

Total length: <1.4 m.

It is found in the eastern Pacific Ocean, from the Gulf of California to Peru, at depths from the surface to 40 m.

carcharhinus coatesi

(Carcharhinus coatesi)

Coates's shark

Акула Коутса

Total length: 70–88 cm.

It is found off northern Australia, from Shark Bay in Western Australia to Fraser Island in Queensland, and possibly also off the coast of New Guinea, at depths from the surface to 123 m.

carcharhinus dussumieri

(Carcharhinus dussumieri)

Whitecheek Shark

Сіра акула білощока

Total length: 1–1.2 m.

It is native to the Indo-Pacific Ocean, where it is found on continental shelves and inshore slopes around islands down to about 170 m. Its range extends from the Arabian Sea and Persian Gulf to Java, Indonesia, Japan, and Australia.

carcharhinus falciformis

(Carcharhinus falciformis)

Silky Shark

Акула шовкова

Total length: 2–3.5 m.
Weight: <346 kg.

It is one of the most abundant sharks in the pelagic zone, and can be found around the world in tropical waters. Primarily an inhabitant of the open ocean, the silky shark is most common from the surface to a depth of 200 m, but may dive to 500 m or more.

carcharhinus fitzroyensis

(Carcharhinus fitzroyensis)

Creek Whaler

Сіра акула північноавстралійська

Total length: 1–1.5 m.

Its range is restricted to northern Australia, between Gladstone in central Queensland and Cape Cuvier in Western Australia. It inhabits estuaries and inshore waters from the intertidal zone to depths of at least 40 m.

carcharhinus galapagensis

(Carcharhinus galapagensis)

Galápagos Shark

Акула галапагоська

Total length: 3–3.7 m.
Weight: ≈195 kg.

It is commonly found in tropical waters near islands in all oceans, at depths of 1–286 m. The largest concentrations occur near archipelagos or island groups, particularly around Bermuda, the Galápagos, Madeira, Cape Verde, the Marshall and Mariana Islands, the Tuamotu Archipelago, and Hawaii. It is also found near the Baja California Peninsula, along the coasts of Guatemala and Colombia, off Portugal and Spain, and near eastern Australia.

carcharhinus hemiodon

(Carcharhinus hemiodon)

Pondicherry Shark

Акула Пондіччері

Total length: 1.5–2 m.

It was once found throughout Indo-Pacific coastal waters from the Gulf of Oman to New Guinea, and is known to enter fresh water. Most of the known specimens were collected from India, with more specimens from the Gulf of Oman, Borneo, and Java.

carcharhinus humani

(Carcharhinus humani)

Human's Whaler

Сіра акула Г’юмана

Total length: ≈84.4 cm.

It inhabits the western Indian Ocean near the Socotra Islands, off Kuwait, Mozambique, and South Africa, at depths of 36–43 m.

carcharhinus isodon

(Carcharhinus isodon)

Finetooth Shark

Сіра акула рівнозуба

Total length: 1.6–1.9 m.

In North American waters, it is found from North Carolina to the northern Gulf of Mexico. In Central and South America, it occurs off Trinidad and Guyana, is infrequently recorded in the Caribbean Sea, and is found off southern Brazil from São Paulo to Santa Catarina. It inhabits extremely shallow waters, no deeper than 20 m.

carcharhinus leiodon

(Carcharhinus leiodon)

Smoothtooth Blacktip Shark

Сіра акула гладенькозуба

Total length: <1.3 m.

It is known only from the type specimen caught from the Gulf of Aden, off eastern Yemen, and a handful of additional specimens caught from the Persian Gulf, off Kuwait.

carcharhinus leucas

(Carcharhinus leucas)

Bull Shark

Акула-бик

Total length: 2.25–3.5 m.
Weight: 95–130 kg.

It is commonly found worldwide in coastal areas of warm oceans, and also occurs in rivers and lakes, occasionally entering saltwater and freshwater streams if they are deep enough, at depths of 0–150 m. It has traveled 4,000 km up the Amazon River. It also lives in freshwater Lake Nicaragua, in the Ganges and Brahmaputra Rivers in Eastern India and adjoining Bangladesh.

carcharhinus limbatus

(Carcharhinus limbatus)

Common Blacktip Shark

Акула чорнопера

Total length: 1.5–2.8 m.
Weight: <123 kg.

It is common to coastal tropical and subtropical waters around the world, including brackish habitats. It is found in water less than 30 m deep over continental and insular shelves, though it may dive to 64 m.

carcharhinus longimanus

(Carcharhinus longimanus)

Oceanic Whitetip Shark

Акула довгокрила

Total length: 1.5–4 m.
Weight: 20–60 kg.

It inhabits the pelagic zone of tropical and warm-temperate seas. It is found worldwide between 45°N and 43°S. It lives in deep, open океans in waters warmer than 18 °C, spending most of its time in the upper ocean layers down to about 150 m.

carcharhinus macloti

(Carcharhinus macloti)

Hardnose Shark

Сіра акула твердолоба

Total length: 80–110 cm.

It is found from Kenya to Myanmar in the Indian Ocean, including Sri Lanka and the Andaman Islands. In the Pacific Ocean, it is found from Vietnam to Taiwan and southern Japan, in Indonesia, and off New Guinea and northern Australia. It is usually found in shallow, inshore waters, but has been reported to a depth of 170 m.

carcharhinus melanopterus

(Carcharhinus melanopterus)

Blacktip Reef Shark

Рифова акула чорнопера

Total length: <1.8 m.
Weight: <13.6 kg.

It is found throughout nearshore waters of the tropical and subtropical Indo-Pacific, where it inhabits coral reefs and prefers shallow, inshore habitats. It has been reported from depths of up to 75 m, but is usually found in waters only a few meters deep.

carcharhinus obscurus

(Carcharhinus obscurus)

Dusky Shark

Акула смуглява

Total length: 3.2–4.2 m.
Weight: 160–347 kg.

It occurs, albeit discontinuously, in tropical and warm-temperate continental seas worldwide. It is typically found at depths of 10–80 m, but may occasionally descend to as deep as 400 m.

carcharhinus obsoletus

(Carcharhinus obsoletus)

Lost Shark

Акула загублена

Total length: <43.3 cm.

Only three specimens of this species are known, found in Borneo, Vietnam, and Thailand, all of which are over 80 years old.

carcharhinus oxyrhynchus

(Carcharhinus oxyrhynchus)

Daggernose Shark

Сіра акула гостроноса

Total length: 1.4–2.4 m.
Weight: ≈13 kg.

It inhabits shallow tropical waters off northeastern South America, from Trinidad to northern Brazil, favoring muddy habitats such as mangroves, estuaries, and river mouths. It occurs at depths of 4–40 m.

carcharhinus perezii

(Carcharhinus perezii)

Caribbean Reef Shark

Карибська рифова акула

Total length: 2–3 m.
Weight: <70 kg.

It is found in the tropical waters of the western Atlantic Ocean from Florida to Brazil, and is the most commonly encountered reef shark in the Caribbean Sea. It is common in water shallower than 30 m, but has been known to dive to 378 m.

carcharhinus plumbeus

(Carcharhinus plumbeus)

Sandbar Shark

Акула пісочна

Total length: 1.8–3 m.
Weight: 80–118 kg.

It is found in tropical to temperate waters of the Atlantic Ocean and the Indo-Pacific. It commonly occurs over muddy or sandy bottoms in shallow coastal habitats such as bays, estuaries, harbors, and river mouths, but it is also found in intertidal zones and deeper waters exceeding 200 m.

carcharhinus porosus

(Carcharhinus porosus)

Smalltail Shark

Дрібнохвоста сіра акула

Total length: 0.9–1.5 m.

It is found in the western Atlantic Ocean, from the northern Gulf of Mexico to southern Brazil. It inhabits shallow waters close to shore, particularly over muddy bottoms around estuaries. It occurs at depths of up to 35 m.

carcharhinus sealei

(Carcharhinus sealei)

Blackspot Shark

Сіра акула Сейла

Total length: 70–100 cm.

It is found in the tropical Indo–West Pacific Ocean between latitudes 24°N and 30°S, occurring on continental shelves and in shallow waters around islands from the surface to depths of about 40 m.

carcharhinus signatus

(Carcharhinus signatus)

Night Shark

Атлантична нічна акула

Total length: 2–2.8 m.
Weight: <67.7 kg.

It is found along the outer continental shelves and upper continental slopes of the Atlantic Ocean, from the U.S. state of Massachusetts to Argentina in the west, including the Gulf of Mexico and the Caribbean Sea, and from Senegal to northern Namibia in the east. It occurs at depths of 50–600 m.

carcharhinus sorrah

(Carcharhinus sorrah)

Spot-tail Shark

Сіра акула плямохвоста

Total length: 1.1–1.6 m.
Weight: <28 kg.

It is found in the tropical Indo-Pacific on continental and insular shelves commonly to a depth around 73 m. Its range extends from the East African coast, Madagascar, and the Red Sea to India, Malaysia, China, the Philippines, and northern Australia.

carcharhinus tilstoni

(Carcharhinus tilstoni)

Australian Blacktip Shark

Австралійська чорнопера акула

Total length: 1.5–2 m.
Weight: <52 kg.

It is endemic to northern and eastern Australia. Favoring the upper and middle parts of the water column, it can be found from the intertidal zone to a depth of 50 m.

carcharhinus tjutjot

(Carcharhinus tjutjot)

Indonesian Whaler

Сіра акула індонезійська

Total length: 70–100 cm.

It is found in the western Pacific, ranging from Indonesia and Taiwan to Borneo. It prefers to live in depths ranging from the surface to 170 m and in demersal inshore habitats.

glyphis glyphis

(Glyphis glyphis)

Speartooth Shark

Річкова акула звичайна

Total length: 1.75–2.6 m.

It inhabits coastal marine waters and tidal reaches of large tropical rivers in northern Australia and New Guinea. It is found in near-shore marine waters, favoring highly turbid environments over a wide range of salinities.

glyphis garricki

(Glyphis garricki)

New Guinea River Shark

Річкова акула північна

Total length: <2.5 m.

It found in scattered tidal rivers and associated coastal waters in northern Australia and in Papua New Guinea. It inhabits areas with poor visibility, soft bottoms, and large tides, with immature sharks ranging into fresh and brackish water.

glyphis gangeticus

(Glyphis gangeticus)

Ganges River Shark

Річкова акула гангська

Total length: 1.78–2 m.

It is found in rivers of eastern and north-eastern India, particularly the Hooghly River in West Bengal, and the Ganges, Brahmaputra, and Mahanadi rivers in Bihar, Assam, and Odisha, respectively. It is typically found in the middle to lower reaches of these rivers.

lamiopsis temminckii

(Lamiopsis temminckii)

Broadfin Shark

Широкопера акула звичайна

Total length: 1.2–1.7 m.
Weight: <52.4 kg.

It is found in the Indian and Pacific Oceans, at depths of up to 50 m. Its distribution is sporadic, with records from off the coasts of Pakistan, India, Indonesia, Myanmar, China, and Sarawak.

lamiopsis tephrodes

(Lamiopsis tephrodes)

Borneo Broadfin Shark

Широкопера акула борнейська

Total length: <1.4 m.
Weight: <13.5 kg.

It is found in the Indo–West Pacific, from Thailand, Indonesia, and Malaysia, and is thought to occur more widely throughout the Indo-Malay Archipelago to southern China, at depths of down to 50 m.

loxodon macrorhinus

(Loxodon macrorhinus)

Sliteye Shark

Акула вузькоока

Total length: <95 cm.

It is found in the tropical waters of the Indo-West Pacific, from the Red Sea and East Africa to Indonesia, north to Japan and south to Australia, between latitudes 34°N and 30°S, at depths of 7–100 m.

prionace glauca

(Prionace glauca)

Blue Shark

Акула блакитна

Total length: 1.82–2.82 m.
Weight: 27–55 kg.

It is an oceanic and epipelagic shark found worldwide in deep temperate and tropical waters from the surface to about 350 m.

negaprion brevirostris

(Negaprion brevirostris)

Lemon Shark

Лимонна акула

Total length: 2.4–3.4 m.
Weight: 90–183.7 kg.

It is found from New Jersey to southern Brazil in the tropical western Atlantic Ocean and also occurs off the coast of West Africa in the south-eastern Atlantic, including around Cape Verde. In addition, it has been recorded in the eastern Pacific from southern Baja California to Ecuador. It occurs at depths of 0–92 m.

negaprion fronto

(Negaprion fronto)

Pacific Lemon Shark

Лимонна акула тихоокеанська

Total length: ≈3.68 m.

It is endemic to the eastern Pacific, where it is found from the Gulf of California to Peru, from the surface to a depth of 92 m.

negaprion acutidens

(Negaprion acutidens)

Sicklefin Lemon Shark

Лимонна акула гострозуба

Total length: 3–3.8 m.

Its range extends from South Africa to the Red Sea (including Mauritius, the Seychelles, and Madagascar), continuing eastward along the coast of the Indian subcontinent to Southeast Asia, and extending as far north as Taiwan and the Philippines and as far south as New Guinea and northern Australia. It is also found around numerous Pacific islands. It occurs at depths of 0–92 m.

rhizoprionodon terraenova

(Rhizoprionodon terraenovae)

Atlantic Sharpnose Shark

Довгоноса акула атлантична

Total length: 70–110 cm.
Weight: <7.3 kg.

It is found in the subtropical waters of the north-western Atlantic Ocean, between latitudes 18°N and 43°N, at depths of 0–280 m.

rhizoprionodon acutus

(Rhizoprionodon acutus)

Milk Shark

Акула молочна

Total length: 1.1–1.78 cm.
Weight: <22 kg.

It is found in the eastern Atlantic Ocean (from Mauritania to Angola, as well as around Madeira), in the Indian Ocean (from South Africa and Madagascar northward to the Arabian Peninsula and eastward to South and Southeast Asia), and in the Pacific Ocean (from China and southern Japan to New Guinea and northern Australia). It occurs close to shore, from the surf zone to depths of 200 m.

rhizoprionodon lalandii

(Rhizoprionodon lalandii)

Brazilian Sharpnose Shark

Довгоноса акула бразильська

Total length: 78–80 cm.

It is found in the tropical waters of the western Atlantic Ocean between latitudes 13° N and 33° S, at depths between 3 and 70 m.

rhizoprionodon longurio

(Rhizoprionodon longurio)

Pacific Sharpnose Shark

Довгоноса акула тихоокеанська

Total length: 96–154 cm.

It inhabits the coastal waters of the eastern tropical Pacific Ocean at depths of up to 27 m and can be found from southern California to Peru.

rhizoprionodon oligolinx

(Rhizoprionodon oligolinx)

Grey Sharpnose Shark

Довгоноса акула сіра

Total length: 55–70 cm.

It is found in the tropical waters of the Indo-West Pacific Oceans, between latitudes 30° N and 18° S, from the surface to a depth of 36 m.

rhizoprionodon porosus

(Rhizoprionodon porosus)

Caribbean Sharpnose Shark

Довгоноса акула карибська

Total length: 80–110 cm.

It is found in the tropical waters of the western Atlantic Ocean and the Caribbean, between latitudes 28° N and 40° S, from the surface to a depth of 500 m.

rhizoprionodon taylori

(Rhizoprionodon taylori)

Australian Sharpnose Shark

Довгоноса акула австралійська

Total length: 50–69,5 cm.

It is found in the tropical waters of the western Pacific Ocean off Papua New Guinea and northern Australia, between latitudes 8°N and 28°S, from the surface to a depth of 110 m.

triaenodon obesus

(Triaenodon obesus)

Whitetip Reef Shark

Рифова акула білопера

Total length: ≈1.6 m.
Weight: <18.3 kg.

It is found on Indo-Pacific coral reefs, occurring as far west as South Africa and as far east as Central America. It is typically found on or near the bottom in clear water at depths of 8–40 m.

scoliodon laticaudus

(Scoliodon laticaudus)

Spadenose Shark

Акула лопатоноса

Total length: 74–100 cm.

It is found in the western Indo-Pacific from Tanzania to South and Southeast Asia, as far east as Java and Borneo and as far north as Taiwan and Japan. It is typically found close to the coast in water 10–13 m deep.

scoliodon macrorhynchos

(Scoliodon macrorhynchos)

Pacific Spadenose Shark

Акула лопатоноса тихоокеанська

Total length: <71 cm.
Weight: <2.2 kg.

It is found in the western Pacific Ocean, from the coast of Malaysia to southern Japan, and is occasionally recorded from the Bay of Bengal. It occurs at depths of up to 10–15 m.

nasolamia velox

(Nasolamia velox)

Whitenose Shark

Акула сіра білоноса

Total length: <1.5 cm.

It it is found in the tropical waters of the eastern Pacific Ocean between latitudes 31° N and 18° S, at depths of 15–200 m.